High-density mapping of the MHC identifies a shared role for HLA-DRB1∗01:03 in inflammatory bowel diseases and heterozygous advantage in ulcerative colitis

Philippe Goyette, Gabrielle Boucher, Dermot Mallon, Eva Ellinghaus, Luke Jostins, Hailiang Huang, Stephan Ripke, Elena S. Gusareva, Vito Annese, Stephen L. Hauser, Jorge R. Oksenberg, Ingo Thomsen, Stephen Leslie, Mark J. Daly, Kristel Van Steen, Richard H. Duerr, Jeffrey C. Barrett, Dermot P.B. McGovern, L. Philip Schumm, James A. TraherneMary N. Carrington, Vasilis Kosmoliaptsis, Tom H. Karlsen, Andre Franke, John D. Rioux, International Inflammatory Bowel Disease Genetics Consortium (IIBDGC), Peter A. Bampton

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